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+[[!meta title="The transposon Galileo generates natural chromosomal inversions in Drosophila by ectopic recombination."]]
+[[!tag Drosophila repeat variants]]
+
+Delprat A, Negre B, Puig M, Ruiz A.
+
+PLoS One. 2009 Nov 18;4(11):e7883. doi:10.1371/journal.pone.0007883
+
+The transposon Galileo generates natural chromosomal inversions in Drosophila by ectopic recombination.
+
+[[!pmid 19936241 desc="In _D. buzzatii_, the 2z(3) inversion is flanked by several transposable elements, among which two Galileo repeats (cut-and-paste mechanism, members of the P element family), that “(i) are inserted in opposite orientation; (ii) present exchanged target site duplications; and (iii) are both chimeric”, suggesting ectopic recombination."]]

diff --git a/tags/Drosophila.mdwn b/tags/Drosophila.mdwn
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--- a/tags/Drosophila.mdwn
+++ b/tags/Drosophila.mdwn
@@ -43,4 +43,7 @@ See also [[Muller elements|muller_element]].
 _D. melanogaster_'s development time course at different temperatures was fitted
 to the Arrhenius law in [[Crapse and coll., 2021|biblio/34414660]].
 
+Ectopic recombination of a Galileo element may have caused a recent large-scale
+inversion in _D. buzzati_ ([[Delprat and coll, 2009|biblio/19936241]]).
+
 [[!inline pages="tagged(Drosophila)" limit=0]]

diff --git a/tags/variants.mdwn b/tags/variants.mdwn
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--- a/tags/variants.mdwn
+++ b/tags/variants.mdwn
@@ -35,6 +35,9 @@ vicinity.  [[Donelly and coll., 2010|biblio/20116045]] gave a radically lower
 time estimate of 13,600 to 108,400 years, but an ancient origin was again
 supported by [[Steinberg and coll in 2012|biblio/22751100]].
 
+Ectopic recombination of a Galileo element may have caused a recent large-scale
+inversion in _D. buzzati_ ([[Delprat and coll, 2009|biblio/19936241]]).
+
 ### Software
 
  - _NanoSV_ ([[Cretu Stancu and coll., 2017|biblio/29109544]]) uses nanopore long