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+[[!meta title="The evolutionary history of placodes: a molecular genetic investigation of the larvacean urochordate Oikopleura dioica."]]
+[[!tag Oikopleura olfaction]]
+
+Bassham S, Postlethwait JH.
+
+Development. 2005 Oct;132(19):4259-72. doi:10.1242/dev.01973
+
+The evolutionary history of placodes: a molecular genetic investigation of the larvacean urochordate _Oikopleura dioica_.
+
+[[!pmid 16120641 desc="Animals fed with algal concentrates.  _O. dioica_ has one orthologue of _eyes absent_, _Eya_, and one orthologue of the vertebrate _Pitx_ genes, with multiple promoters and alternative splice sites, and lacking an OAR domain.  It has three _sine oculis_ orthologues, _Six1/2_, _Six3/6a_ and _Six3/6b.  They are not clustered like vertebrate genes, but the two _Six3/6_ share an intron position in their homeodomain.  Expression pattern of these genes support the hypothesis that _O. dioica_ has a placode that is homologous to the vertebrate olfactory placode."]]

diff --git a/tags/Oikopleura.mdwn b/tags/Oikopleura.mdwn
index 0bc923fee0f7f737a5a3222559d87cf4ad80ea66..a81e4dc685aca745b6819d841a413e2c396c9d13 100644 (file)
--- a/tags/Oikopleura.mdwn
+++ b/tags/Oikopleura.mdwn
@@ -258,6 +258,8 @@ Genes and pathways
  - _O. dioica_ has the miRNA machinery and some miRNAs such as _let-7a_ were detected.  36 % of
    the miRNAs had a length of 22 nt in [[Fu, Adamski and Thompson, 2008|biblio/18339653]].
  - 6 FGF genes were detected by Oulion, Bertrand and Escriva ([[2012|biblio/22919541]]).
+ - _Eya_, _Pitx_, _Six1/2_, _Six3/6a_ and _Six3/6b_, with expression patterns supporting
+   homology of larvacean and vertebrate placodes ([[Bassham and Postlethwait, 2005|biblio/16120641]]).
 
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