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+[[!meta title="Tunicates illuminate the enigmatic evolution of chordate metallothioneins by gene gains and losses, independent modular expansions and functional convergences."]]
+[[!tag Oikopleura]]
+
+Calatayud S, Garcia-Risco M, Palacios Ò, Capdevila M, Cañestro C, Albalat R.
+
+Mol Biol Evol. 2021 Jun 19:msab184. doi:10.1093/molbev/msab184
+
+Tunicates illuminate the enigmatic evolution of chordate metallothioneins by gene gains and losses, independent modular expansions and functional convergences.
+
+[[!pmid 34146103 desc="Ancestral two-domain organisation found in Cephalochordates and Vertebrates.  Ascidians appear to have lost one domain and larvaceans proteins ressemble no other."]]

diff --git a/tags/Oikopleura.mdwn b/tags/Oikopleura.mdwn
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--- a/tags/Oikopleura.mdwn
+++ b/tags/Oikopleura.mdwn
@@ -234,6 +234,8 @@ Genes and pathways
    Tolstenkov and Glover 2019|biblio/30905687]]).
  - Piwi ([[Henriet and coll., 2015|biblio/25779047]]).
  - Metallothioneins _OdMT1_ and _OdMT2_ [[Calatayud and coll., 2018|biblio/30284576]].
+   Their cysteins are organised in a way that is not found in other branches of the
+   evolutionary tree [Calatayud and coll., 2021|biblio/34146103].
  - 2 NUMB genes were found; both are closer to Vertebrate NUMB than to Vertebrate NUMB-Like
    ([[Confalonieri and coll., 2019|biblio/31451549]]).
  - Genes related to thyroid functions (_vWFL_, _Nkx2-1_, _FoxE_, _TPO_ and _Duox_)