Et ensuite, des râmen

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+[[!meta title="Pyrococcus genome comparison evidences chromosome shuffling-driven evolution."]]
+[[!tag archaea variants]]
+
+Zivanovic Y, Lopez P, Philippe H, Forterre P.
+
+Nucleic Acids Res. 2002 May 1;30(9):1902-10. doi:10.1093/nar/30.9.1902
+
+Pyrococcus genome comparison evidences chromosome shuffling-driven evolution. 
+
+[[!pmid 11972326 desc="“the extent of rearrangement that occurred after the
+three species diverged is much lower between _P. abyssi_ and _P. horikoshii_
+than between these two species and _P. furiosus_.”  “[In _P. furiosus_, the
+distribution of IS elements] is clearly non-random, since 16 of them are
+located near shuffled segments boundaries, five are in completely scrambled
+regions and only two are found within an undisturbed segment.”  “the AT3 skew
+was less upset than the word skew (here GGTT), which was itself, as expected,
+less perturbed than the gene orientation skew”.  Coding genes were weakly
+colinear with the replication direction in _P. abissi_ and P. horikoshii_,
+especially for highly expressed genes, but this correlation was less evident or
+absent for _P. furiosus_."]]

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@@ -65,6 +65,10 @@ this phenomenon was confirmed by [[D'Iorio and Dewar, 2022|biblio/36261227]].
 The state of the art in bacterial genomics in 2000 was reviewed by
 [[Hughes|biblio/11380986]].
 
+A mixture of X-shaped dot-plot pattern and scrambling was reported by
+[[Zivanovic and coll. (2002)|biblio/11972326]] in a study of _Pyrococcus_
+genomes.
+
 ### Mechanism
 
 _Staggered breaks_ ([[Ranz and coll., 2007|bilbio/17550304]]) caused