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+[[!meta title="Generation of a widespread Drosophila inversion by a transposable element."]]
+[[!tag variants]]
+
+Cáceres M, Ranz JM, Barbadilla A, Long M, Ruiz A.
+
+Science. 1999 Jul 16;285(5426):415-8. doi:10.1126/science.285.5426.415
+
+Generation of a widespread Drosophila inversion by a transposable element.
+
+[[!pmid 10411506 desc="Target Site Duplication (TSD) sites were exchanged through the recombination of two transposons inserted in opposite directions."]]

diff --git a/tags/variants.mdwn b/tags/variants.mdwn
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+++ b/tags/variants.mdwn
@@ -38,6 +38,10 @@ supported by [[Steinberg and coll in 2012|biblio/22751100]].
 Ectopic recombination of a Galileo element may have caused a recent large-scale
 inversion in _D. buzzati_ ([[Delprat and coll, 2009|biblio/19936241]]).
 
+Flanking an inversion in _D. buzzati_ between two _Galileo_ elements, the
+exchange of Target-Site Duplication sequences was observed by Cáceres and
+coll., [[(1999|biblio/10411506]]). 
+
 [[Kent and coll., 2003|biblio/14500911]] reported 2 inversions per Mbp in
 human/mouse comparisons, median length 814.